Eusociality Tools & Calculators
Free, hands-on tools for the science of social life — work out genetic relatedness, test Hamilton's rule, project a colony's growth, estimate a hive's population, map a foraging range, and decode the waggle dance. Educational estimates only.
Coefficient of Relatedness Calculator
Read off Wright's coefficient of relatedness (r) for any relationship — from identical twins to first cousins — including the r = 0.75 shared by haplodiploid sisters in bees, ants, and wasps that helps explain eusociality
Hamilton's Rule Calculator
Test rB > C, the equation at the heart of social evolution: enter relatedness, benefit, and cost to see whether kin selection favours an act of altruism and its inclusive-fitness effect
Colony Growth Calculator
Project a colony's worker force over the season from the queen's laying rate, survival to adulthood, development time, and worker lifespan — with the month-by-month curve and steady-state size
Honeybee Hive Population Estimator
Turn a hive inspection into a head count with the beekeeper's frame-count method — adult bees from bee-covered frames plus the sealed brood waiting to emerge
Colony Foraging Range Calculator
Estimate the area a colony can work as a disc of π r² around the nest, in km² and hectares, with species presets and an optional overlap check for colonies packed at a given density
Waggle Dance Decoder
Decode a honeybee's waggle dance — waggle-run duration gives the distance to the food, and the dance angle plus the sun's azimuth gives a compass bearing to the source